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Abstract A long‐standing question is how felsic continental crust is differentiated from its mafic parent mantle magmas. One currently proposed fundamental mechanism is lithospheric foundering and loss of dense material into the mantle. A type locality is the young extinct arc forming the Sierra Nevada, California. Here, we image a distinct anisotropic shear layer below the crust‐mantle boundary using receiver functions. The sense of shear is consistent with west‐ to southwestward removal of lithosphere. The shear signal is strongest in the southern Sierra, where lithospheric foundering was proposed to have concluded several million years ago, and is deeper and less pronounced in the central Sierra, where ongoing lithospheric foundering is corroborated by a band of unusually deep (40+ km) seismicity along the western foothills. Our observations provide progressive snapshots of a lithospheric foundering process spanning several million years and hundreds of kilometers, illuminating a fundamental differentiation process by which continents are built.